Dr. Igor Khorozyan

Research Scientist

Curriculum Vitae:

from 2016: Research scientist at the Department of Conservation Biology, Georg-August-Universität Göttingen, Germany
2012-2016: Postdoctoral scientist at the Department of Conservation Biology, Georg-August-Universität Göttingen, Germany
2011-2012: Lecturer at American University of Armenia and WWF expert, Yerevan, Armenia
2007-2011: Project coordinator at World Wide Fund for Nature (WWF), Yerevan, Armenia
2006-2007: Manager at Foundation for Biodiversity Conservation in Armenian Highland, Yerevan, Armenia
1999-2007: Freelance researcher, Yerevan, Armenia
1997-2003: Research fellow at Centre for Ecological Studies, National Academy of Sciences, Yerevan, Armenia

Expert, trainer and consultant for different national and international projects

Research interests - large mammalian carnivores:

Non-invasive techniques of wildlife research and monitoring
Species taxonomy, biogeography and evolution
Population ecology and in-situ conservation
Effect of natural and man-caused factors on populations
Managing co-existence between wildlife and humans
Conservation approaches (biodiversity monitoring, establishment and strengthening of protected areas)


Wildlife assessment
International nature conservation
Statistics for field biologists
Ecology and conservation of threatened species
Environmental field study (certificate programme)
Environmental science and conservation

Current projects:

    Completed projects (2012-2019):

    Major publications:

    Ahmadpour M., Shokri S., Varasteh-Moradi H., Waltert M., Khorozyan I., Soofi M. 2021. The occurrence of the rare corsac fox (Vulpes corsac) in Iran is mainly determined by prey presence and land use. Journal of Arid Environments 189: 104475. https://doi.org/10.1016/j.jaridenv.2021.104475

    Farhadinia M.S., Rostro-Garcia S., Feng L., Kamler J.F., Spalton A., Shevtsova E., Khorozyan I., Al-Duais M., Ge J. and Macdonald D.W. 2021. Big cats in borderlands: challenges and implications for transboundary conservation of Asian leopards. Oryx 55: 452-460. https://doi.org/10.1017/S0030605319000693DOI:10.1017/S0030605319000693

    Khorozyan I., Ananian V., Malkhasyan A. 2021. No longer regionally extinct: a review of Pallas’s cat Otocolobus manul records from the Caucasus with a new record from Armenia (Mammalia: Felidae). Zoology in the Middle East 67: 12-18. https://doi.org/10.1080/09397140.2020.1865663

    Khorozyan I., Waltert M. 2021. A global view on evidence-based effectiveness of interventions used to protect livestock from wild cats. Conservation Science and Practice 3:e317. https://doi.org/10.1111/csp2.317

    Filla M., Lama R.P., Ghale T.R., Signer J., Filla T., Aryal R.R., Heurich M., Waltert M., Balkenhol N., Khorozyan I. 2021. In the shadows of snow leopards and the Himalayas: density and habitat selection of blue sheep in Manang, Nepal. Ecology and Evolution 11: 108-122. https://doi.org/10.1002/ece3.6959

    Bruns A., Waltert M. and Khorozyan I. 2020. The effectiveness of livestock protection measures against wolves (Canis lupus) and implications for their co-existence with humans. Global Ecology and Conservation 21: e00868. https://doi.org/10.1016/j.gecco.2019.e00868

    Khorozyan I. 2020. A comparison of common metrics used to quantify the effectiveness of conservation interventions. PeerJ 8: e9873. http://doi.org/10.7717/peerj.9873

    Khorozyan I., Ghoddousi S., Soufi M., Soofi M. and Waltert M. 2020. Studded leather collars are very effective in protecting cattle from leopard (Panthera pardus) attacks. Ecological Solutions and Evidence 1: e12013. https://doi.org/10.1002/2688-8319.12013 Media Coverage: [The Applied Ecologist]

    Khorozyan I., Waltert M. 2020. Variation and conservation implications of the effectiveness of anti‑bear interventions. Scientific Reports 10: 15341. https://doi.org/10.1038/s41598-020-72343-6

    Ashrafzadeh M.R., Naghipour A.A., Haidarian M. and Khorozyan I. 2019. Modeling the response of an endangered flagship predator to climate change in Iran. Mammal Research 64: 39-51. https://doi.org/10.1007/s13364-018-0384-y

    Bleyhl B., Arakelyan M., Askerov E., Bluhm H., Gavashelishvili A., Ghoddousi A., Heidelberg A., Khorozyan I., Malkhasyan A., Manvelyan K., Masoud M., Moqanaki E., Radeloff V.C., Soofi M., Weinberg P., Zazanashvili N. and Kümmerle T. 2019. Assessing niche overlap between domestic and threatened wild sheep to identify conservation priority areas. Diversity and Distributions 25: 129-141. https://doi.org/10.1111/ddi.12839

    Ghoddousi A., Soofi M., Hamidi A. Kh., Ashayeri S., Egli L., Ghoddousi S., Speicher J., Khorozyan I., Kiabi B.H. and Waltert M. 2019. The decline of ungulate populations in Iranian protected areas calls for urgent action against poaching. Oryx 53: 151-158. https://doi.org/10.1017/S003060531600154X

    Khorozyan I. and Waltert M. 2019. A framework of most effective practices in protecting human assets from predators. Human Dimensions of Wildlife 24: 380-394. https://doi.org/10.1080/10871209.2019.1619883

    Khorozyan I. and Waltert M. 2019. How long do anti-predator interventions remain effective? Patterns, thresholds and uncertainty. Royal Society Open Science 6: 190826. https://doi.org/10.1098/rsos.190826

    Soofi M., Ghoddousi A., Zeppenfeld T., Shokri S., Soufi M., Egli L., Jafari A., Ahmadpour M., Qashqaei A., Ghadirian T., Filla M., Kiabi B., Balkenhol N., Waltert M. and Khorozyan I. 2019 Assessing the relationship between the intensity of illegal hunting of ungulates, wild prey occurrence and livestock depredation rate by large carnivores. Journal of Applied Ecology 56: 365-374. https://doi.org/10.1111/1365-2664.13266

    Khorozyan I., Ghoddousi S., Soufi M., Soofi M. and Waltert M. 2018. Cattle selectivity by leopards suggests ways to mitigate human-leopard conflict. Ecology and Evolution 8: 8011–8018. DOI:10.1002/ece3.4351

    Meinecke L., Soofi M., Riechers M., Khorozyan I., Hosseini H., Schwarze S. and Waltert M. 2018. Crop variety and prey richness affect spatial patterns of human-wildlife conflicts in Iran’s Hyrcanian forests. Journal for Nature Conservation 43: 165-172. DOI:10.1016/j.jnc.2018.04.005

    Sogbohossou E.A, Kassa B.D., Waltert M. and Khorozyan I. 2018. Spatio-temporal niche partitioning between the African lion (Panthera leo leo) and spotted hyena (Crocuta crocuta) in western African savannas. European Journal of Wildlife Research 64: 1. DOI:10.1007/s10344-017-1159-5

    Soofi M., Ghoddousi A., Zeppenfeld T., Shokri S., Soufi M., Jafari A., Ahmadpour M., Qashqaei A.T., Egli L., Ghadirian T., Chahartagh N.R., Zehzad B., Kiabi B.H., Khorozyan I., Balkenhol N. and Waltert M. 2018. Livestock grazing in protected areas and its effects on large mammals in the Hyrcanian forest, Iran. Biological Conservation 217: 377–382. DOI:10.1016/j.biocon.2017.11.020

    Filla M., Premier J., Magg N., Dupke C., Khorozyan I., Waltert M., Bufka L. and Heurich M. 2017. Habitat selection by Eurasian lynx (Lynx lynx) is primarily driven by avoidance of human activity during day and prey availability during night. Ecology and Evolution 7: 6367-6381. DOI:10.1002/ece3.3204

    Ghoddousi A., Egli L, Soofi M., Khorozyan I. and Waltert M. 2017. After sanctions: the urge to upgrade and integrate conservation in Iran. Frontiers in Ecology and the Environment 15: 9-10. DOI:10.1002/fee.1452

    Ghoddousi A., Soofi M., Hamidi A. Kh., Lumetsberger T., Egli L., Ashayeri S., Khorozyan I., Kiabi B.H. and Waltert M. 2017. When pork is not on the menu: Assessing trophic competition between large carnivores and poachers. Biological Conservation 209: 223-229. DOI:10.1016/j.biocon.2017.02.032

    Khorozyan I., Lumetsberger T., Ghoddousi A., Soofi M. and Waltert M. 2017. Global patterns of biomass models describing prey consumption by big cats. Mammal Review 47: 124-132. DOI:10.1111/mam.12084

    Khorozyan I., Soofi M., Soufi M., Hamidi A. Kh., Ghoddousi A. and Waltert M. 2017. Effects of shepherds and dogs on livestock depredation by leopards (Panthera pardus) in north-eastern Iran. PeerJ 5: e3049. DOI:10.7717/peerj.3049

    Lumetsberger T., Ghoddousi A., Appel A., Khorozyan I., Waltert M. and Kiffner C. 2017. Re-evaluating models for estimating prey consumption by leopards. Journal of Zoology 302: 201-210. DOI:10.1111/jzo.12449

    Soofi M., Ghoddousi A., Egli L., Khorozyan I. and Waltert M. 2017. Letter to editor: incorrect identification of study species. Comparative Clinical Pathology 26: 9-10. DOI:10.1007/s00580-016-2371-5

    Soofi M., Ghoddousi A., Hamidi A.Kh., Ghasemi B., Egli L., Voinopol-Sassu A-J., Kiabi B.H., Balkenhol N., Khorozyan I. and Waltert M. 2017. Precision and reliability of indirect population assessments for the Caspian red deer (Cervus elaphus maral). Wildlife Biology 1: wlb.00230. DOI:10.2981/wlb.00230

    Ghoddousi A., Hamidi A.Kh., Soofi M., Khorozyan I., Kiabi B.H. and Waltert M. 2016. Effects of ranger stations on predator and prey distribution and abundance in an Iranian steppe landscape. Animal Conservation 19: 273-280. DOI:10.1111/acv.12240

    Ghoddousi A., Soofi M., Hamidi A.Kh., Lumetsberger T., Egli L., Khorozyan I., Kiabi B.H. and Waltert M. 2016. Assessing the role of livestock in big cat prey choice using spatiotemporal availability patterns. PLoS One 11: e0153439. DOI: 10.1371/journal.pone.0153439

    Proulx G., Abramov A.V., Adams I., Jennings A.P., Khorozyan I., Rosalino L.M., Santos-Reis M., Veron G. and Do Linh San E. 2016. World distribution and status of badgers - a review. In: Badgers: Systematics, Biology, Conservation and Research Techniques (Proulx G., Do Linh San E., eds.). Sherwood Park, Alpha Wildlife Publications: 31-116.

    Khorozyan I., Ghoddousi A., Soofi M. and Waltert M. 2015. Big cats kill more livestock when wild prey reaches a minimum threshold. Biological Conservation 192: 268-275. DOI: 10.1016/j.biocon.2015.09.031

    Khorozyan I., Soofi M., Hamidi A.K., Ghoddousi A. and Waltert M. 2015. Dissatisfaction with veterinary services is associated with leopard (Panthera pardus) predation on domestic animals. PLoS One 10 (6): e0129221

    Khorozyan I., Soofi M., Ghoddousi A., Hamidi A.K. and Waltert M. 2015. The relationship between climate, diseases of domestic animals and human-carnivore conflicts. Basic and Applied Ecology 16: 703-713. DOI:10.1016/j.baae.2015.07.001

    Khorozyan I. 2014. Morphological variation and sexual dimorphism of the common leopard (Panthera pardus) in the Middle East and their implications for species taxonomy and conservation. Mammalian Biology 79: 398-405. DOI:10.1016/j.mambio.2014.07.004

    Khorozyan I., Stanton D., Mohammed M., Al-Ra'il W. and Pittet M. 2014. Patterns of co-existence between humans and mammals in Yemen: some species thrive while others are nearly extinct. Biodiversity and Conservation 23: 1995-2013. DOI:10.1007/s10531-014-0700-z.

    Khorozyan I., Malkhasyan A. and Murtskhvaladze M. 2011. The striped hyaena Hyaena hyaena (Hyaenidae, Carnivora) rediscovered in Armenia. Folia Zoologica 60 (3): 253-261.

    Ghoddousi A., Hamidi Kh. A., Ghadirian T., Ashayeri D. and Khorozyan I. 2010. The status of the endangered Persian leopard Panthera pardus saxicolor in Bamu National Park, Iran. Oryx 4 (4): 551-557.

    Khorozyan I.G., Malkhasyan A.G., Asmaryan S.G. and Abramov A.V. 2010. Using geographical mapping and occupancy modeling to study the distribution of the critically endangered leopard (Panthera pardus) population in Armenia. In: Spatial Complexity, Informatics, and Wildlife Conservation (Cushman S., Huettmann F., eds.). Tokyo, Springer Publishers: 331-347.

    Khorozyan I.G., Weinberg P.I. and Malkhasyan A.G. 2009. Conservation strategy for Armenian mouflon (Ovis [orientalis] gmelini Blyth) and bezoar goat (Capra aegagrus Erxleben) in Armenia. In: Status and Protection of Globally Threatened Species in the Caucasus (Zazanashvili N., Mallon D., eds.). Tbilisi, CEPF, WWF: 37-46.

    Khorozyan I.G., Malkhasyan A.G. and Abramov A.V. 2008. Presence-absence surveys of prey and their use in predicting leopard (Panthera pardus) densities: a case study from Armenia. Integrative Zoology 3 (4): 322-332.

    Khorozyan I.G., Cazon A., Malkhasyan A.G. and Abramov A.V. 2007. Using thin-layer chromatography of fecal bile acids to study the leopard (Panthera pardus ciscaucasica) population. Biology Bulletin 34 (4): 361-366.

    Khorozyan I.G. and Abramov A.V. 2007. The leopard, Panthera pardus (Carnivora: Felidae), and its resilience to human pressure in the Caucasus. Zoology in the Middle East 41: 11-24.

    Khorozyan I.G., Baryshnikov G.F. and Abramov A.V. 2006. Taxonomic status of the leopard, Panthera pardus (Carnivora, Felidae) in the Caucasus and adjacent areas. Russian Journal of Theriology 5 (1): 41-52.

    Khorozyan I., Malkhasyan A. and Asmaryan S. 2005. The Persian leopard prowls its way to survival. Endangered Species Update 22: 51-60.

    Khorozyan I. 2003. Habitat preferences by the endangered Persian leopard (Panthera pardus saxicolor Pocock, 1927) in Armenia. Zoology in the Middle East 30: 25-36.