Research interests


  • Evolution and systematics of seed plants, particularly the gymnosperms.

  • Carboniferous environments and plants.

  • The origin, taphonomy, and fossil record of amber, in particular the global phenomena of ‘amber bursts’, and the plant inclusions enclosed within amber.

  • Understanding the interactions of species in ‘amber forests’.

  • Resin induction and production in modern plants, particularly the gymnosperms.

  • Anatomically preserved fossil plants and reconstructing the fragments back in to a whole-plant concept.

  • Niche development and the geological record.

  • Land plant recovery after extinction events, particularly the Triassic floral recovery.

  • Origin of the modern flora.