Simon S, Letsch H, Bank S, Buckley TR, Donath A, Liu S, Machida R, Meusemann K, Misof B, Podsiadlowski L, Zhou X, Wipfler B, Bradler S 2019. Old World and New World Phasmatodea: Phylogenomics resolve the evolutionary history of stick and leaf insects. Frontiers in Ecology and Evolution 7: 345.

Büscher TH, Grohmann C, Bradler S, Gorb SN 2019. Tarsal attachment pads in Phasmatodea (Hexapoda: Insecta). Zoologica 164: 1–94.

Glaw F, Hawlitschek O, Dunz A, Goldberg J, Bradler S 2019. When giant stick insects play with colors: Molecular phylogeny of the Achriopterini and description of two new splendid species (Phasmatodea: Achrioptera) from Madagascar. Frontiers in Ecology and Evolution 7: 105.


Robertson JA, Bradler S, Whiting MF 2018. Evolution of oviposition techniques in stick and leaf insects (Phasmatodea). Frontiers in Ecology and Evolution 6: 216.

Bradler S, Buckley TR 2018. Biodiversity of Phasmatodea. In: Insect Biodiversity: Science and Society Vol 2 – eds Foottit RG, Adler PH (John Wiley & Sons): 281–313.

Büscher T, Buckley TR, Grohmann C, Gorb SN, Bradler S 2018. The evolution of tarsal adhesive microstructures in stick and leaf insects (Phasmatodea). Frontiers in Ecology and Evolution 6, 69.


Leubner F, Wipfler B, Bradler S 2017. Comparative morphology of the orthopteran thorax with a discussion of phylogenetically relevant characters. Insect Systematics and Diversity 1: 29–47.

Leubner F, Bradler S, Wipfler B 2017. The thoracic morphology of the wingless dune cricket Comicus calcaris (Orthoptera: Schizodactylidae): Novel apomorphic characters for the group and adaptations to sand desert environments. Arthropod Structure and Development 46: 449–461.


Shelomi M, Danchin EGJ, Heckel D, Wipfler B, Bradler S, Zhou X, Pauchet Y 2016. Horizontal gene transfer of pectinases from bacteria preceded the diversification of stick and leaf insects. Scientific Reports 6: 26388.

Leubner F, Hörnschemeyer T, Bradler S 2016. The thorax of the cave cricket Troglophilus neglectus: anatomical adaptations in an ancient wingless insect lineage (Orthoptera: Rhaphidophoridae). BMC Evolutionary Biology 16: 39.


Archibald SB, Bradler S 2015. Stem-group stick insects (Phasmatodea) in the early Eocene at McAbee, BC, Canada, and Republic, Washington, USA. The Canadian Entomologist 147: 744-753.

Bradler S 2015. The Phasmatodea Tree of Life: Surprising facts and open questions in the evolution of stick and leaf insects. Entomologie heute 27: 1-23.

Bradler S, Cliquennois N, Buckley TR 2015. Single origin of Mascarene stick insects: ancient radiation on sunken islands? BMC Evolutionary Biology 15: 196.

Stumpner A, Stritih N, Mai O, Bradler S 2015 Diversity of Orthoptera in the south-western Karst-region of Slovenia with notes on acoustics and species identification. Acta Entomologica Slovenica 23: 5-20.

Goldberg J, Bresseel J, Constant J, Kneubühler B, Leubner F, Michalik P, Bradler S 2015. Extreme convergence in egg-laying strategy across insect orders. Scientific Reports 5: 7825.


Buckley TR, Myers SS, Bradler S 2014. Revision of the stick insect genus Clitarchus Stål (Phasmatodea: Phasmatidae): new synonymies and two new species from northern New Zealand. Zootaxa 3900: 451-482.

Wang M, Béthoux O, Bradler S, Jacques F, Cui Y, Ren D 2014. Under cover at pre-angiosperm times: a cloaked phasmatodean insect from the Early Cretaceous Jehol biota. PLOS ONE 9: e91290.

Bradler S, Robertson JA, Whiting MF 2014. A molecular phylogeny of Phasmatodea with emphasis on Necrosciinae, the most species-rich subfamily of stick insects. Systematic Entomology 39: 205-222.


Dönitz J, Grossmann D, Schild I, Bradler S, Prpic N.-M, Bucher G 2013. TrOn: An anatomical ontology for the beetle Tribolium castaneum. PLOS ONE 8: e70695.

Hörnschemeyer T, Haug JT, Béthoux O, Beutel RG, Charbonnier S, Hegna TA, Koch M, Rust J, Wedmann S, Bradler S, Willmann R 2013. Is Strudiella a Devonian insect? Nature 494: E3-E4.


Wirmer A, Bradler S, Heinrich R 2012. Homology of the insect corpora allata and vertebrate adenohypophysis? Arthropod Structure & Development 41: 409-417.

Friedemann K, Wipfler B, Bradler S, Beutel RG 2012. On the head morphology of Phyllium and the phylogenetic relationships of Phasmatodea (Insecta). Acta Zoologica 93: 184-199.


Helm C, Treulieb S, Werler K, Bradler S, Klass K-D 2011. The male genitalia of Oxyartes lamellatus. Phasmatodeans do have complex phallic organs (Insecta: Phasmatodea). Zoologischer Anzeiger 250: 223-245.

Bradler S, Buckley TR 2011. Stick insect on unsafe ground: Does a fossil from the Early Eocene of France really link Mesozoic taxa to the extant crown group of Phasmatodea? Systematic Entomology 36: 218-222.


Buckley TR, Bradler S 2010. Tepakiphasma ngatikuri, a new genus and species of stick insect (Phasmatodea) from the Far North of New Zealand. New Zealand Entomologist 33: 118-126.

Buckley TR, Attanayake D, Nylander JAA, Bradler S 2010. The phylogenetic placement and biogeographic origins of the New Zealand stick insects (Phasmatodea). Systematic Entomology 35: 207-225.


Bradler S 2009. Phylogenie der Stab- und Gespenstschrecken (Insecta: Phasmatodea). Species, Phylogeny & Evolution 2: 3-139.

Buckley TR, Attanayake D, Bradler S 2009. Extreme convergence in stick insect evolution: phylogenetic placement of the Lord Howe island tree lobster. Proceedings of the Royal Society London Series B 276: 1055-1062.


Wedmann S, Bradler S, Rust J 2007. The first fossil leaf insect: 47 million years of specialized cryptic morphology and behavior. Proceedings of the National Academy of Sciences of the United States of America 104: 565-569.


Klug R, Bradler S 2006. The pregenital abdominal musculature in phasmids and its implications for the basal phylogeny of Phasmatodea (Insecta: Polyneoptera). Organisms, Diversity & Evolution 6: 171-184.


Bradler S, Whiting MF, Klug R 2003. Basal Diversification and the evolution of wings within stick insects (Phasmatodea). Entomologische Abhandlungen 61: 132-133.

Whiting MF, Bradler S, Maxwell T 2003. Loss and recovery of wings in stick insects. Nature 421: 264-267.


Bradler S 2001. Die australischen Stabschrecken ? die ?Beuteltiere? unter den Phasmatodea? Verhandlungen des Westdeutschen Entomologen Tages 2000: 179-184.


Bradler S 1999. The vomer of Timema Scudder, 1895 (Insecta: Phasmatodea) and its significance for phasmatodean phylogeny. Courier Forschungsinstitut Senckenberg 215: 43-47.


Bradler S 2009. Kommen sie nie zurück? Möglichkeiten einer Re-Evolution. ? In: Elsner N, Fritz H-J, Gradstein R, Reitner J (eds): Evolution ? Zufall und Zwangsläufigkeit der Schöpfung, Wallstein-Verlag: 261-292.

Bradler S 2003. Phasmatodea, Gespenstschrecken. In: Dathe HH (ed): Kaestner, Lehrbuch der speziellen Zoologie, I, 5: 251-261. Gustav Fischer/Spektrum Akademischer Verlag.


Bradler S, Seiler C 2012. Phasmiden ? Biologie, Haltung und Zucht. Verlag Eugen Ulmer, Stuttgart. 175 pp.

Seiler C,Bradler S,Koch R 2006. Fasmidos. Insectos palo e insectos hoja. Hispano Europea, Barcelona. 162 pp.

Seiler C, Bradler S, Koch R 2000. Ratgeber Phasmiden. Bede-Verlag, Ruhmannsfelden. 144 pp.

BOOK REVIEWS (invited)

Bradler S 2012. Book review: In search of the causes of evolution. Grant PR, Grant BR (eds) Princeton University Press 2010. In: Basic and Applied Ecology 13: 639.

Bradler S 2010. Book review: Speciation and patterns of diversity. Butlin RK, Bridle JR, Schluter D (eds). Cambridge University Press 2009. In: Basic and Applied Ecology 11: 469.


Bradler S, Buckley TR 2011. Monophyly and ancient radiation of the Mascarene stick insects. In: Borsch T, Giere P, Hoffmann J, Jahn R, Löhne C, Nordt B, Ohl M (eds): Biosystematics 2011: 66.

Bradler S, Wedmann S 2008. Diversity and paleobiology of Eocene stick and leaf insects (Insecta: Phasmatodea). In: Gradstein SR, Klatt S, Normann F, Weigelt P, Willmann R, Wilson R (eds): Systematics 2008: 174.

Buckley TR, Bradler S 2008: Extreme convergence in stick insect evolution (Phasmatodea): phylogenetic placement of the rarest insect of the world. In: Gradstein SR, Klatt S, Normann F, Weigelt P, Willmann R, Wilson R (eds): Systematics 2008: 46.

Wedmann S, Bradler S, Rust J 2007. The first fossil walking leaf (Insecta: Phasmatodea). An example of evolutionary stasis. In: Hörweg C, Sattmann H (eds): Abstracts from the 9th annual meeting of the GfBS: 113. Organisms, Diversity & Evolution 7: 252.

Bradler S, Mai O 2007. A phylogeny of stick and leaf insects (Insecta: Phasmatodea) with special emphasis on re-evolution events. In: Hörweg C, Sattmann H (eds): Abstracts from the 9th annual meeting of the GfBS: 15. Organisms, Diversity & Evolution 7: 252.

Bradler S 2002. Evolution of secondary ovipositors in stick insects (Insecta: Phasmatodea). Zoology 105, Supplement V (DZG 95.1): 60.

Bradler S 2001. The Australian stick insects - a monophyletic group within the Phasmatodea? Zoology 104, Supplement III (DZG 94.1): 69.

Bradler S 2000. On the systematic position of Agathemera Stål 1875 within the Phasmatodea (Insecta). Zoology 103, Supplement III (DZG 93.1): 99.