Prof. Dr. Andreas Schuldt


  • 2015 Habilitation and venia legendi in Ecology; Leuphana Universität Lüneburg
  • 2010 Doctoral degree in Ecology (Dr. rer. nat.), summa cum laude, Leuphana Universität Lüneburg
  • 2007 Diploma in Biology, Georg-August-Universität Göttingen

Professional Experience:

  • since 2018 Professor and Head of the Department of Forest Nature Conservation, Georg-August-Universität Göttingen
  • 2016-2018 Postdoctoral researcher, German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig
  • 2010-2017 Postdoctoral researcher/fellowship, Leuphana Universität Lüneburg

Awards & Fellowships:

  • 2010-2014 Postdoctoral Fellowship; Leuphana Universität Lüneburg
  • 2010 Best Doctoral Researcher 2009 (3rd place); Leuphana Universität Lüneburg
  • 2007-2010 PhD Scholarship; Leuphana Universität Lüneburg

Editorial Board:

  • Biodiversity and Conservation (Associate Editor)
  • Journal of Plant Ecology (Associate Editor)



Hagge, J., Müller, J., Bässler, C., Brandl, R., Schuldt, A., Thorn, S., Seibold, S. (2024) Change in saproxylic beetle, fungi and bacteria assemblages along horizontal and vertical gradients of sun-exposure in forest. Biological Conservation, 291, 1–10.

Singer, D., Hagge, J., Kamp, J., Hondong, H., Schuldt, A. (2024) Aggregated time-series features boost species-specific differentiation of true and false positives in passive acoustic monitoring of bird assemblages. Remote sensing in Ecology and Conservation.

Vázquez-González, C., Castagneyrol, B., Muiruri, E. W., Barbaro, L., Abdala-Roberts, L., Barsoum, N., Fründ, J., Glynn, C., Jactel, H., McShea, W. J., Mereu, S., Mooney, K. A., Morillas, L., Nock., C. A., Paquette, A., Parker, J. D., Parker, W. C., Roales, J., Scherer-Lorenzen, M., Schuldt, A., Verheyen, K., Weih, M., Yang, B., Koricheva, J. (2024) Tree diversity enhances predation by birds but not by arthropods across climate gradients. Ecology Letters, 27 (5), 1–11.

Wildermuth, B., Hagge, J., Seifert, C.L., Tjaden, R. & Schuldt, A. (2024) Beneficial effects of native broadleaved forests on canopy beetle diversity are not reduced by admixture of non-native conifers. Journal of Applied Ecology, 61, 1000-1014.

Wildermuth, B., Penanhoat, A., Sennhenn-Reulen, H., Matevski, D., Drescher, J., Aubry-Kientz, M., Seidel, D., Schuldt, A. (2024) Canopy structure influences arthropod communities within and beyond tree identity effects: Insights from combining LiDAR data, insecticidal fogging and machine learning regression modelling. Ecological Indicators,

Yi, H., Eisenhauer, N., Austen, J.C, Rebollo, R., Ray, T., Bönisch, E., von Oheimb, G., Fichtner, A., Schuldt, A., Patoine, G., Ferlian, O. (2024) Tree diversity and mycorrhizal type co-determine multitrophic ecosystem functions. Journal of Ecology, 112 (3), 528-546,


Chen, J.-T., Wang, M.-Q., Li, Y., Chesters, D., Luo, A., Zhang, W., Guo, P.-F., Guo, S.-K., Zhou, Q.-S., Ma, K.-P., von Oheimb, G., Kunz, M., Zhang, N.-L., Liu, X.-J., Bruelheide, H., Schuldt, A., Zhu, C.-D. (2023) Functional and phylogenetic relationships link predators to plant diversity via trophic and non-trophic pathways. Proceedings of the Royal Scociety B 290 (1990)

Glatthorn, J., Appleby, S., Balkenhol, N., Kriegel, P., Likulunga, L. E., Lu, J. Z., Matevski, D., Polle, A., Riebl, H., Rivera Pérez, C. A., Scheu, S., Seinsche, A., Schall, P., Schuldt, A., Wingender, S., Ammer, C. (2023) Species diversity of forest-floor biota in non-native Douglas fir stands is similar as in native stands. Ecosphere, 14, 1-16,

Li, Y., Du, Y.-B., Chen, J.-T., Wang, M.-Q., Guo, S.-K., Schuldt, A., Luo, A., Guo, P.-F., Mi, X.-C., Liu, X.-J., Ma, K.-P., Bruelheide, H., Chesters, D., Liu, X., Zhu, C.-D. (2023) Tree dissimilarity determines multi-dimensional beta-diversity of herbivores and carnivores via bottom-up effects. Journal of Animal Ecology 92 (2), 442-453

Li, Y., Schmid, B., Schuldt, A., Li, S., Wang, M.-Q., Fornoff, F., Staab, M., Guo, P.-F., Anttonen, P., Chesters, D., Bruelheide, H., Zhu, C.-D., Ma, K., Liu, X. (2023) Multitrophic arthropod diversity mediates tree diversity effects on primary productivity. Nature Ecology & Evolution

Li, Y., Wang, M.-Q., Chesters, D., Anttonen, P., Guo, S.-K., Guo, P.-F., Chen, J.-T., Ma, K., Bruelheide, H., Schuldt, A., Zhu, C.-D., (2023) Differential impacts on herbivore diversity and scale-dependence of tree diversity in subtropical forests. Journal of Ecology, 111 (3), 666-675

Matevski, D., Foltran, E., Lamersdorf, N., Schuldt, A. (2023) Introduction of non-native Douglas fir reduces leaf damage on beech saplings and mature trees in European beech forests. Ecological Applications 33 (2)

Matevski, D., Schuldt, A. (2023) Non-native Douglas fir promotes epigeal spider density, but has a mixed effect on functional diversity. Biodiversity and Conservation 32, 1233-1250

Schuldt, A., Liu, X., Buscot, F., Bruelheide, H., Erfmeier, A., He, J.-S., Klein, A.-M., Ma, K., Scherer-Lorenzen, M., Schmid, B., Scholten, T., Tang, Z., Trogisch, S., Wirth, C., Wubet, T., Staab, M. (2023) Carbon–biodiversity relationships in a highly diverse subtropical forest. Global Change Biology

Wang, M.-Q., Wen, Z., Ke, J., Chesters, D., Li, Y., Chen, J.-T., Luo, A., Shi, X., Zhou, Q.-S., Liu, X-J., Ma, K., Bruelheide, H., Schuldt, A., Zhu, C.-D., (2023) Tree communities and functional traits determine herbivore compositional turnover. Oecologia, 203, 205-218.

Wildermuth, B., Dönges, C., Matevski, D., Penanhoat, A., Seifert, C. L., Seidel, D., Scheu, S. & Schuldt, A. (2023) Tree species identity, canopy structure and prey availability differentially affect canopy spider diversity and trophic composition. Oecologia, 203, 37-51.

Wildermuth, B., Fardiansah, R., Matevski, D., Lu, J.-Z., Kriegel, P., Scheu, P., Schuldt, A. (2023) Conifers and non-native tree species shift trophic niches of generalist arthropod predators in Central European beech forests. BMC Ecology and Evolution 23,

Wildermuth, B., Seifert, C.L., Husemann, M. & Schuldt, A. (2023) Metabarcoding reveals that mixed forests mitigate negative effects of non-native trees on canopy arthropod diversity. ECOLOGICAL APPLICATIONS, 33: e2921, 1-17.


Anttonen, P., Li, Y., Chesters, D., Davrinche, A., Haider, S., Bruelheide, H., Chen, J.-T., Wang, M.-Q., Ma, K.-P., Zhu, C.-D., Schuldt, A. (2022) Leaf Nutritional Content, Tree Richness, and Season Shape the Caterpillar Functional Trait Composition Hosted by Trees. insects 13,

Huang, Y., Schuldt, A., Hönig, L., Yang, B., Liu, X., Bruelheide, H., Ma, KP., Schmid, B., Niklaus, P. (2022) Effects of enemy exclusion on biodiversity-productivity relationships in a subtropical forest experiment. Journal of Ecology (accepted)

Schuldt, A., Huke, P., Glatthorn, J., Hagge, J., Wildermuth, B., Matevski, D. (2022) Tree mixtures mediate negative effects of introduced tree species on bird taxonomic and functional diversity. Journal of Applied Ecology 59 (12), 3049-3060

Tang, T., Zhang, N., Bongers, F. J., Staab, M., Schuldt, A. Fornoff, F., Lin, H., Cavender-Bares, J., Hipp, A., Li, S., Liang, Y., Han, B., Klein, A.-M., Bruelheide, H., Durka, W., Schmid, B., Ma, K., Liu, X. (2022) Tree species and genetic diversity increase productivity via functional diversity and trophic feedbacks. eLife

Wang, M.-Q., Yan, C., Luo, A., Li, Y., Chesters, D., Qiao, H.-J., Chen, J.-T., Zhou, Q.-S., Ma, K., Bruelheide, H., Schuldt, A., Zhang, Z., Zhu, C.-D. (2022) Phylogenetic relatedness, functional traits, and spatial scale determine herbivore co-occurrence in a subtropical forest. Ecological Monographs 92 (1), 1-16


Glatthorn J, Annighöfer P, Balkenhol N, Leuschner C, Polle A, Scheu S, Schuldt A, Schuldt B, Ammer C (2021) An interdisciplinary framework to describe and evaluate the functioning of forest ecosystems. Basic and Applied Ecology 52, 1-14

Kriegel P, Matevski D, Schuldt A (2021) Monoculture and mixture-planting of non-native Douglas fir alters species composition, but promotes the diversity of ground beetles in a temperate forest system Biodiverstity and Conservation 30, 1479–1499

Li Y, Chesters D, Wang MQ, Wubet T, Schuldt A, Anttonen P, Guo PF, Chen JT, Zhou QS, Zhang N, Ma, KP, Bruelheide H, Wu CS, Zhu CD (2021) Tree diversity and functional leaf traits drive herbivore-associated microbiomes in subtropical China. Ecology and Evolution

Löffler F, Schuldt A, König-Ries B, Bruelheide H, Klan F (2021) A Test Collection for Dataset Retrieval in Biodiversity Research. Research Ideas and Outcomes. Research Ideas and Outcomes

Matevski D, Glatthorn J, Kriegel P, Schuldt A (2021) Non-native Douglas fir (Pseudotsuga menziesii) promotes sentinel prey attack rates in Central European forests. Forest Ecology and Management 489, 119099

Matevski D, Schuldt A (2021) Tree species richness, tree identity and non-native tree proportion affect arboreal spider diversity, abundance and biomass. Forest Ecology and Management, 483, 118775

Rutten G, Hönig L, Braun U, Michalski S, Saadani M, Schwass R, Schuldt A, Bruelheide H (2021) More diverse tree communities promote foliar fungal diversity, but decrease infestation rates per tree species in a subtropical biodiversity experiment. Journal of Ecology 109, 2068-2080

Staab M, Liu X, Assmann T, Bruelheide H, Buscot F, Durka W, Erfmeier A, Klein Alexandra M, Ma K, Michalski S, Wubet T, Schmid B, Schuldt A (2021) Tree phylogenetic diversity structures multitrophic communities. Functional Ecology 35 (2), 521-534

Trogisch S, Liu X, Rutten G, Xue K, Bauhus J, Brose U, Bu W, Cesarz S, Chesters D, Connolly J, Cuid X, Eisenhauer N, Guo L, Haider S, Härdtle W, Kunz M, Ma L, Neumann S, Sang W, Schuldt A, Tang Z, Van Dam N, Oheim G, Wang M, Wang S, Weinhold A, Wirth C, Wubet T, Xu X, Yang B, Zhang N, Zhu C, Ma K, Wang Y, Bruelheide H (2021) The significance of tree-tree interactions for forest ecosystem functioning. Basic and Applied Ecology

Valdés-Correcher E, Moreira X, Augusto L, Barbaro L, Bouget C, Bouriaud O, Branco M, Centenaro G, Csóka G, Damestoy T, Dobrosavljević J, Duduman M-L, Dulaurent A-M, Eötvös CB, Faticov M, Ferrante M, Fürjes-Mikó Á, Galmán A, Gossner MM, Hampe A, Harvey D, Gordon Howe A, Kadiri Y, Kaennel-Dobbertin M, Koricheva J, Kozel A, Kozlov MV, Lövei GL, Lupaștean D, Milanović S, Mrazova A, Opgennoorth L, Pitkänen J-M, Popova A, Popović M, Prinzing A, Queloz V, Roslin T, Sallé A, Sam K, Scherer-Lorenzen M, Schuldt A, Selikhovkin A, Suominen L, Tack AJM, Tahadlova M, Thomas R, Castagneyrol B (2021) Search for top-down and bottom-up drivers of latitudinal trends in insect herbivory in oak trees in Europe. Global Ecology and Biogeography 30, 651-665

Ming-Qiang Wang, Chuan Yan, Arong Luo, Yi Li, Douglas Chesters, Hui-Jie Qiao, Jing-Ting Chen, Qing-Song Zhou, Keping Ma, Helge Bruelheide, Andreas Schuldt, Zhibin Zhang, Chao-Dong Zhu (2021) Phylogenetic relatedness, functional traits, and spatial scale determine herbivore co-occurrence in a subtropical forest. Ecological Monographs 92 (1), 1-16

Zumstein P, Bruelheide H, Fichtner A, Schuldt A, Staab M, Härdtle W, Zhou H, Assmann T (2021) What shapes ground beetle assemblages in a tree species-rich subtropical forest? ZooKeys 1044, 907-927


Wang MQ, Li Y, Chesters D, Bruelheide H, Ma KP, Guo PF, Zhou QS, Staab M, Zhu CD, Schuldt A (2020) Host functional and phylogenetic composition rather than host diversity structure plant-herbivore networks. Molecular Ecology.

Staab M, Schuldt A (2020) The Influence of Tree Diversity on Natural Enemies - a Review of the “Enemies” Hypothesis in Forests. Current Forestry Reports.

Fichtner A, Schnabel F, Bruelheide H, Kunz M, Mausolf K, Schuldt A, Härdtle W, von Oheimb G (2020) Neighbourhood diversity mitigates drought impacts on tree growth. Journal of Ecology.

Hülsmann M, Boutaud E, Buse J, Frank K, Schuldt A, Assmann T (2020) Habitat continuity matters: Ancient woodlands tend to have higher biomass and catching rate of dung beetles, mainly driven by one large species. Acta Oecologica 102: 103501.


Eisenhauer N, Schielzeth H, Barnes AD, Barry K, Bonn A, Brose U, Bruelheide H, Buchmann N, Buscot F, Ebeling A, Ferlian O, Freschet GT, Giling DP, Hättenschwiler S, Hillebrand H, Hines J, Isbell F, Koller-France E, König-Ries B, de Kroon H, Meyer ST, Milcu A, Müller J, Nock CA, Petermann JS, Roscher C, Scherber C, Scherer-Lorenzen M, Schmid B, Schnitzer SA, Schuldt A, Tscharntke T, Türke M, van Dam NM, van der Plas F, Vogel A, Wagg C, Wardle DA, Weigelt A, Weisser WW, Wirth C, Jochum M (2019) A multitrophic perspective on biodiversity–ecosystem functioning research. Advances in Ecological Research 61: 1-54 DOI: 10.1016/bs.aecr.2019.06.001

Wang MQ, Li Y, Chesters D, Anttonen P, Bruelheide H, Chen JT, Durka W, Guo PF, Härdtle W, Ma KP, Michalski SG, Schmid B, von Oheimb G, Wu CS, Zhang NL, Zhou QS, Schuldt A, Zhu CD (2019) Multiple components of plant diversity loss determine herbivore phylogenetic diversity in a subtropical forest experiment. Journal of Ecology 107: 2697-2712 DOI: 10.1111/1365-2745.13273

Schuldt A, Ebeling A, Kunz M, Staab M, Guimarães-Steinicke C, Bachmann D, Buchmann N, Durka W, Fichtner A, Fornoff F, Härdtle W, Hertzog L, Klein AM, Roscher C, Schaller J, von Oheimb G, Weigelt A, Weisser W, Wirth C, Zhang J, Bruelheide H, Eisenhauer N (2019) Multiple plant diversity components drive consumer communities across ecosystems. Nature Communications 10: 1460. DOI: 10.1038/s41467-019-09448-8

Hülsmann M, Boutaud E, Buse J, Schuldt A, Assmann T (2019) Land-use legacy and tree age in continuous woodlands: weak effects on overall ground beetle assemblages, but strong effects on two threatened species. Journal of Insect Conservation 23: 623-633. DOI: 10.1007/s10841-019-00156-8

Nolte D, Boutaud E, Kotze DJ, Schuldt A, Assmann T (2019) Habitat specialization, distribution range size and body size drive extinction risk in carabid beetles. Biodiversity and Conservation 28: 1267-1283


Huang Y, Chen Y, Castro-Izaguirre N, Baruffol M, Brezzi M, Lang A, Li Y, Härdtle W, von Oheimb G, Yang X, Liu X, Pei K, Both S, Yang B, Eichenberg D, Assmann T, Bauhus J, Behrens T, Buscot F, Chen X-Y, Chesters D, Ding B-Y, Durka W, Erfmeier A, Fang J, Fischer M, Guo L-D, Guo D, Gutknecht JLM, He J-S, He C-L, Hector A, Hönig L, Hu R-Y, Klein A-M, Kühn P, Liang Y, Li S, Michalski S, Scherer-Lorenzen M, Schmidt K, Scholten T, Schuldt A, Shi X, Tan M-Z, Tang Z, Trogisch S, Wang Z, Welk E, Wirth C, Wubet T, Xiang W, Yu M, Yu X-D, Zhang J, Zhang S, Zhang N, Zhou H-Z, Zhu C-D, Zhu L, Bruelheide H, Ma K, Niklaus PA, Schmid B Impacts of species richness on productivity in a large-scale subtropical forest experiment. Science 362: 80–83

Schuldt A, Assmann T, Brezzi M, Buscot F, Eichenberg D, Gutknecht J, Härdtle W, He JS, Klein AM, Kühn P, Liu X, Ma K, Niklaus PA, Pietsch KA, Purahong W, Scherer-Lorenzen M, Schmid B, Scholten T, Staab M, Tang Z, Trogisch S, Oheimb G von, Wirth C, Wubet T, Zhu CD, Bruelheide H (2018) Biodiversity across trophic levels drives multifunctionality in highly diverse forests. Nature Communications 9:2989. DOI: 10.1038/s41467-018-05421-z

Binkenstein J, Klein AM, Assmann T, Buscot F, Erfmeier A, Ma K, Pietsch K, Schmidt K, Scholten T, Wubet T, Bruelheide H, Schuldt A, Staab M (2018) Trophic guilds respond differently to changing elevation in a subtropical forest. Ecography 41: 1013-1023

Grossman JJ, Vanhellemont M, Barsoum N, Bauhus J, Bruelheide H, Castagneyrol B, Cavender-Bares J, Eisenhauer N, Ferlian O, Gravel D, Hector A, Jactel H, Kreft H, Mereu S, Messier C, Muys B, Nock C, Paquette A, Parker J, Perring MP, Ponette Q, Reich PB, Schuldt A, Staab M, Weih M, Zemp DC, Scherer-Lorenzen M, Verheyen K (2018) Synthesis and future research directions linking tree diversity to growth, survival, and damage in a global network of tree diversity experiments. Environmental and Experimental Botany 152: 68-89

Yang B, Li B, He Y, Zhang L, Bruelheide H, Schuldt A (2018) Tree diversity has contrasting effects on predation rates by birds and arthropods on three broadleaved, subtropical tree species. Ecological Research 33:205-212


Schuldt A, Fornoff F, Bruelheide H, Klein AM, Staab M (2017) Tree species richness attenuates the positive relationship between mutualistic ant-hemipteran interactions and leaf chewer herbivory. Proceedings of the Royal Society B 20171489. DOI: 10.1098/rspb.2017.1489

Trogisch S, Schuldt A, Bauhus J, Blum J, Both S, Buscot F, Castro-Izaguirre N, Chesters D, Durka W, Eichenberg D, Erfmeier A, Fischer M, Geißler C, Germany M, Goebes P, Gutknecht J, Hahn C, Haider S, Härdtle W, He JS, Hector A, Hönig L, Huang Y, Klein AM, Kühn P, Kunz M, Leppert K, Li Y, Liu X, Niklaus P, Pei Z, Pietsch K, Prinz R, Proß T, Scherer-Lorenzen M, Schmidt K, Scholten T, Seitz S, Song Z, Staab M, von Oheimb G, Weißbecker C, Welk E, Wirth C, Wubet T, Yang B, Zhu CD, Schmid B, Ma K, Bruelheide H: Towards a methodical framework for comprehensively assessing forest multifunctionality. Ecology & Evolution 7: 10652–10674

Schuldt A, Hönig L, Li Y, Fichtner A, Härdtle W, von Oheimb G, Welk E, Bruelheide H (2017) Herbivore and pathogen effects on tree growth are additive, but mediated by tree diversity and plant traits. Ecology & Evolution 7: 7462-7474

Nolte D, Schuldt A, Gossner MM, Ulrich W, Assmann T (2017) Functional traits drive ground beetle community structures in Central European forests: implications for conservation. Biological Conservation 213: 5-12

Schuldt A, Bruelheide H, Assmann T, Buscot F, Erfmeier A, Klein AM, Ma K, Scholten T, Staab M, Wirth C, Zhang J, Wubet T (2017) Belowground top-down and aboveground bottom-up effects structure multitrophic community relationships in a biodiverse forest. Scientific Reports 7: 4222. DOI:10.1038/s41598-017-04619-3

O’Brien MJ, Brezzi M, Schuldt A, Zhang J, Ma K, Schmid B, Niklaus PA (2017) Tree diversity drives diversity of arthropod herbivores but successional stage mediates detritivores. Ecology & Evolution 7: 8753–8760

Zhang J, Bruelheide H, Chen X, Eichenberg D, Kröber W, Xu X, Xu L, Schuldt A (2017) Tree diversity promotes generalist herbivore community patterns in a young subtropical forest experiment. Oecologia 183: 455-467

Brezzi M, Schmid B, Niklaus P, Schuldt A (2017) Tree diversity increases levels of herbivore damage in a subtropical forest canopy: evidence for dietary mixing by arthropods? Journal of Plant Ecology 10: 13-27


Schuldt A, Wubet T, Buscot F, Staab M, Assmann T, Böhnke-Kammerlander M, Both S, Erfmeier A, Klein AM, Ma K, Pietsch K, Schultze S, Wirth C, Zhang J, Zumstein P, Bruelheide H (2015) Multitrophic diversity in a biodiverse forest is highly nonlinear across spatial scales. Nature Communications 6: 10169. DOI: 10.1038/ncomms10169

Schuldt A, Bruelheide H, Härdtle W, Assmann T, Li Y, Ma K, von Oheimb G, Zhang J (2015) Early positive effects of tree species richness on herbivory in a large-scale forest biodiversity experiment influence tree growth. Journal of Ecology 103: 563-571

Seitz S, Goebes P, Zumstein P, Schuldt A, Assmann T, Kühn P, Niklaus P, Scholten T (2015) The influence of leaf litter diversity and soil fauna on initial soil erosion in subtropical forests. Earth Surface Processes and Landforms 40: 1439-1447

Schuldt A, Staab M (2015) Tree species richness strengthens relationships between ants and the functional composition of spider assemblages in a highly diverse forest. Biotropica 47: 339-346

Schuldt A, Baruffol M, Bruelheide H, Chen S, Chi X, Wall M, Assmann T (2014) Woody plant phylogenetic diversity mediates bottom-up control of arthropod biomass in species-rich forests. Oecologia 176: 171-182

Staab M, Schuldt A, Assmann T, Bruelheide H, Klein AM (2014) Ant community structure during forest succession in a subtropical forest in South-East China. Acta Oecologica 61: 32-40

Staab M, Schuldt A, Assmann T, Klein AM (2014) Tree diversity promotes predator but not omnivore ants in a subtropical Chinese forest. Ecological Entomology 39: 637-647

Härdtle W, Niemeyer T, Fichtner A, Li Y, Ries C, Schuldt A, Walmsley D. von Oheimb G (2014) Climate imprints on tree-ring δ15N signatures of sessile oak (Quercus petraea Liebl.) on soils with contrasting water availability. Ecological Indicators 45: 45-50

Homburg K, Homburg N, Schäfer F, Schuldt A, Assmann T (2014) – A dynamic online database of ground beetle species traits (Coleoptera, Carabidae). Insect Conservation and Diversity 7: 195-205


Schuldt A, Scherer-Lorenzen M (2014) Non-native tree species (Pseudotsuga menziesii) strongly decreases predator biomass and abundance in mixed-species plantations of a tree diversity experiment. Forest Ecology and Management 327: 10-17

Schuldt A, Assmann T Bruelheide H, Durka W, Eichenberg D, Härdtle W, Kröber W, Michalski SG, Purschke O (2014) Functional and phylogenetic diversity of woody plants drive herbivory in a highly diverse forest. New Phytologist 202: 864-873

Bruelheide H, Assmann T, Bauhus J, Both S, Buscot F, Chen XY, Durka W, Erfmeier A, Gutknecht JLM, Guo D, Guo LD, Härdtle W, He JS, Klein AM, Kühn P, Michalski S, Nadrowski K, Niklaus P, Pei K, Scherer-Lorenzen M, Scholten T, Schuldt A, Seidler G, Trogisch S, von Oheimb G, Welk E, Wirth C, Wubet T, Yang X, Yu M, Zhang S, Zhou H, Fischer M, Ma K, Schmid B (2014) Designing forest biodiversity experiments: general considerations illustrated by a new large experiment in subtropical China. Methods in Ecology and Evolution 5: 74-89

Schuldt A, Bruelheide H, Durka W, Michalski SG, Purschke O, Assmann T (2014) Tree diversity promotes functional dissimilarity and maintains functional richness despite species loss in predator assemblages. Oecologia 174: 533-543 Assmann T, Drees C, Härdtle W, Klein A, Schuldt A, von Oheimb G (2014) Ökosystem und Biodiversität. In: Heinrichs H, Michelsen G (eds) Nachhaltigkeitswissenschaften. Springer, Berlin, Heidelberg: 147-174 2013

Härdtle W, Niemeyer T, Assmann T, Baiboks S, Fichtner A, Friedrich U, Lang A, Neuwirth B, Pfister L, Ries C, Schuldt A, Simon N, von Oheimb G (2013) Long-term trends in tree-ring width and isotope signatures (δ13C, δ15N) of Fagus sylvatica L. on soils with contrasting water supply. Ecosystems 16: 1413-1428

Härdtle W, Niemeyer T, Assmann T, Aulinger A, Fichtner A, Lang A, Leuschner C, Neuwirth B, Pfister L, Quante M, Ries C, Schuldt A, von Oheimb G (2013) Climatic responses of tree-ring width and δ13C signatures of sessile oak (Quercus petraea Liebl.) on soils with contrasting water supply. Plant Ecology 214: 1147-1156

Schuldt A, Assmann T, Schaefer M (2013) Scale-dependent diversity patterns affect spider assemblages of two contrasting forest ecosystems. Acta Oecologica 49: 17-22

Krause RH, von Wehrden H, Härdtle W, Schuldt A, Assmann T (2013) Stenotopy and eurytopy – distribution models as a tool for estimating niche overlap in two spider species, Trochosa terricola and Eresus kollari (Araneae: Lycosidae / Eresidae). Entomologia Generalis 34: 169-180

Homburg K, Schuldt A, Drees C, Assmann T (2013) Broad-scale geographic patterns in body size and hind wing development of western Palaearctic ground beetles (Coleoptera: Carabidae). Ecography 36: 166-177

Assmann T, Buse J, Dieker P, Eggers B, Drees C, Harry I, Homburg K, Krause H, Matern A, Schuldt A, Taboada A (2013) Historisch alte Waldstandorte: Bedeutung und Wert von Lebensraumtradition für Tiere. In: "Natura 2000 und Wald - Lebensraumtypen, Erhaltungszustand, Management", Naturschutz und Biologische Vielfalt 131: 65-81


Schuldt A, Bruelheide H, Durka W, Eichenberg D, Fischer M, Kröber W, Härdtle W, Ma K, Michalski S, Palm WU, Schmid B, Welk E, Zhou H, Assmann T (2012) Plant traits affecting herbivory on tree recruits in highly diverse subtropical forests. Ecology Letters 15: 732-739

Schuldt A, Bruelheide H, Härdtle W, Assmann T (2012) Predator assemblage structure and temporal variability of species richness and abundance in forests of high tree diversity. Biotropica 44: 793-800


Schuldt A, Both S, Bruelheide H, Härdtle W, Schmid B, Zhou H, Assmann T (2011) Predator diversity and abundance provide little support for the enemies hypothesis in forests of high tree diversity. PLoS ONE 6: e22905

Schuldt A, Assmann T (2011) Patterns and hotspots of carabid beetle diversity in the Palaearctic – insights from a hyperdiverse invertebrate taxon. In Zachos F, Habel JC (eds) Biodiversity Hotspots. Springer, Heidelberg

Bruelheide H, Böhnke M, Both S, Fang T, Assmann T, Baruffol M, Bauhus J, Buscot F, Chen XY, Ding BY, Durka W, Erfmeier A, Fischer M, Geißler C, Guo D, Guo LD, Härdtle W, He JS, Hector A, Kröber W, Kühn P, Lang AC, Nadrowski K, Pei K, Scherer-Lorenzen M, Shi X, Scholten T, Schuldt A, Trogisch S, von Oheimb G, Welk E, Wirth C, Wu YT, Yang X, Zeng X, Zhang S, Zhou H, Ma K, Schmid B (2011) Community assembly during secondary forest succession in a Chinese subtropical forest. Ecological Monographs 81: 25-41

Schuldt A, Assmann T (2011) Belowground carabid beetle diversity in the western Palaearctic – effects of history and climate on range-restricted taxa (Coleoptera: Carabidae). ZooKeys 100: 461-474

Drees C, Brandmayr P, Buse J, Dieker P, Gürlich S, Habel J, Harry I, Härdtle W, Matern A, Meyer H, Pizzolotto R, Quante M, Schäfer K, Schuldt A, Taboada A, Assmann T (2011) Poleward range expansion without a southern contraction in the ground beetle Agonum viridicupreum (Coleoptera, Carabidae). ZooKeys 100: 333-352

Assmann T, Drees C, Matern A, Schuldt A (2011) Eucamaragnathus desenderi, a new ground beetle species from Africa (Coleoptera, Carabidae). ZooKeys 100: 37-46


Schuldt A, Assmann T (2010) Invertebrate diversity and national responsibility for species conservation across Europe – A multi-taxon approach. Biological Conservation 143: 2747-2756

Lang AC, Härdtle W, Bruelheide H, Geißler C, Nadrowski K, Schuldt A, Yu M, von Oheimb G (2010) Growth responses of tree individuals to neighbourhood competition in a species rich slope forest of subtropical China. Forest Ecology and Management 260: 1708-1715

Schuldt A, Baruffol M, Böhnke M, Bruelheide H, Härdtle W, Lang AC, Nadrowski K, von Oheimb G, Voigt W, Zhou, H, Assmann T (2010) Tree diversity promotes insect herbivory in subtropical forests of southeast China. Journal of Ecology 98: 917-926

Assmann T, Casale A, Drees C, Habel JC, Matern A, Schuldt A (2010) The dark side of relict species biology. Cave animals as ancient lineages. In: Habel JC, Assmann T (eds) Relict Species – Phylogeography and Conservation Biology. Springer, Heidelberg


Schuldt A, Wang Z, Zhou H, Assmann T (2009) Integrating highly diverse invertebrates into broad-scale analyses of cross-taxon congruence across the Palaearctic. Ecography 32: 1019-1030

Schuldt A, Assmann T (2009) Environmental and historical effects on richness and endemism patterns of carabid beetles in the Western Palaearctic. Ecography 32: 705-714


Finch OD, Blick T, Schuldt A (2008) Macroecological patterns of spider species richness across Europe. Biodiversity and Conservation 17: 2849-2868

Assmann T, Buse J, Drees C, Habel J, Härdtle W, Matern A, von Oheimb G, Schuldt A, Wrase DW (2008) From Latreille to DNA systematics - towards a modern synthesis for carabidology. In: Penev L, Erwin T, Assmann T (eds) Back to the roots and back to the future. Towards a new synthesis between taxonomic, ecological and biogeographical approaches in carabidology. Pensoft, Sofia, Moscow

Schuldt A, Fahrenholz N, Brauns M, Migge-Kleian S, Platner C, Schaefer M (2008) Communities of ground-living spiders in deciduous forests: Does tree species diversity matter? Biodiversity and Conservation 17: 1267-1284